I am wondering if dogs only howl due to fear or as learned behaviour.
Are there other reasons a dog would howl?
Are lonelines or other emotions a factor?
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Howling is an ancestral trait in dogs as they're descended from wolves, so yes, there could be many reasons for this behavior and it's usually completely natural, it's just a type of dog's vocalization like barking. It's hard to tell exact reason without context and details because there could be as many potential reasons for howling as there could be for barking. It could happen for example as a response to specific sounds, for attention and communication. Loneliness or emotional distress mentioned by you could be valid reasons as well.